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Cover: Measurement of muscle length changes during steady swimming in the shortfin mako shark Isurus oxyrinchus (B) reveals that shortening of the deep red muscle (red trace) is out of phase with surrounding white muscle (white trace), but coincident with local midline curvature about 20% more posterior on the body (see J. M. Donley, R. E. Shadwick, C. A. Sepulveda, P. Konstantinidis and S. Gemballa, pp. 2377-2387), compared to passive swimming (A) where the shortening is coincident. Morphological investigations identify the long tendinous linkages that project red muscle forces from the mid-body to the caudal region. Most prominent are the hypaxial lateral tendons, which span approximately 20% of the body length (inset).

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