- Received September 5, 2016.
- Accepted March 14, 2017.
- Published online March 17, 2017.
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- 1School of Life Sciences, Arizona State University, P.O. Box 874501, Tempe, AZ 85287-4501, USA
- 2Ecology and Evolutionary Biology, University of Colorado, Boulder
- 3US Department of Agriculture, Arid-Land Agricultural Research Center
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