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This Article Summary Full Text (PDF) Alert me when this article is cited Alert me if a correction is posted Services Email this article to a friend Similar articles in this journal Similar articles in PubMed Alert me to new issues of the journal Download to citation manager Citing Articles Citing Articles via HighWire Citing Articles via Google Scholar Google Scholar Articles by Loudon, C. Articles by Koehl, M. A. Search for Related Content PubMed PubMed Citation Articles by Loudon, C. Articles by Koehl, M. A.