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Fig. 5. Strokes A and B generate almost the same amount of force. The only
difference occurs near the end of the stroke when the wing reverses its pitch.
In A the leading edge remains the same, and in B it switches. The wing
pitching in stroke A can be facilitated by wing inertia and aerodynamic torque
as the wing decelerates (Berman and Wang,
2007; Bergou et al.,
2007).