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Fig. 5. A summary of how visual information controls cuttlefish camouflage. The
animal detects local visual features, which include edge and depth
information, and from these relatively low levels then classifies the
background. For example, on the basis of whether it is a continuous surface,
or made of discrete objects such as pebbles, and on the spatial scale of the
pattern/objects. This classification determines the primary weightings
(W1–3) of the components of the coloration pattern. Image contrast (and
perhaps other low-level measures) then modulate the strength of the pattern.
It is unlikely that the classification of `background type' is categorical, in
the sense that an image has to be of one type or another, and this is why the
animal is able to vary the relative levels of expression of the 40 or so
chromatic components independently.
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