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Fig. 6. Planar optode images illustrating the behavioural response of a buried
sandeel (arrow) to a progressive decrease of the oxygen saturation in the
overlying water. The upper row of greyscale photos shows sandeel position and
the lower row the corresponding oxygen images. In the illustrated case, the
fish stayed completely submerged until oxygen saturation in the above water
had declined to 
40%, whereafter its head emerged from the sediment.
Despite increasingly lower levels of oxygen in the above water, the fish
stayed in this position until it finally fully emerged after approximately 140
min at 5–8% air saturation. The time (min) is time elapsed from the
onset of the experiment.
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