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Fig. 4. Deflection of lesser upper- and underwing coverts during a mid-flight gust
response over a sea cliff. (A–D) Covert feather deployment and
(E–H) covert feather retraction. Interval between images: 20 ms between
rows; 320 ms between columns. The coverts remain in approximately the same
deflected position between frames D and E. An animation of the complete
sequence is provided in supplementary material Movie 4.