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Fig. 3. Wing kinematics differ depending upon a bird's wing design and flight
speed. (A) Birds with pointed, high-aspect ratio wings such as the pigeon
Columba livia transition from tip-reversal upstrokes during slow
flight to feathered upstrokes at intermediate speeds and a swept-wing upstroke
during fast flight. (B) Birds with rounded, low-aspect ratio wings such as the
black-billed magpie Pica hudsonica use a flexed upstroke at all
flight speeds. Shown are wingtip (filled circles) and wrist (open circles)
paths in dorsal and lateral view (from
Tobalske and Dial, 1996).