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Fig. 2. Spatial properties of six natural substrates (S1–S6) used in this
study. Left column, images of substrates; right column, rock size
distributions. Rock size scales (i.e. x-axis) of plots are kept the
same to show shift towards the left (i.e. smaller size) of mean rock size. Red
vertical line on the very left of each plot is truncated sand grain size. S1,
rock substrate without any sand; S2, rock substrate with half a cup of the
0 sand spread over the substrate evenly; S3, one cup of sand; S4, two
cups of sand; S5, four cups of sand; S6, full sand.