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Fig. 4. Respiratory behavior in freely behaving, yoked and operantly conditioned
snails. Pneumostome activity was measured over three 45 min training sessions
for (A) freely behaving animals (N=18), (B) yoked animals
(N=17) and (C) operantly conditioned animals (N=31). Values
are means ± s.e.m. for number of pneumostome openings. Animals
operantly conditioned show a significant decrease in pneumostome openings
across sessions. The other two treatment groups show no difference in
pneumostome openings over the training time course. 1, Training session 1; 2,
Training session 2; MT, Memory Test; NS, not significant.
*P<0.05 compared with performance during Training
session 1 within each treatment group.