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Fig. 6. Mean peak GRFs of single legs in geckos climbing (D–F) and running on level ground (A–C). (A) On a level, normal GRFs were always positive. (B) Geckos running over level ground used the forelegs to produce only deceleratory forces, while hindlegs first produced deceleratory forces during the first part of each step, and then produce acceleratory forces during the second part of each step. (C) All four legs pushed laterally away from the midline of the body such that the left legs produced forces that pushed the gecko to the right, while the right legs produced forces that pushed the gecko to the left. (D) In climbing geckos, forelegs produced forces that pushed the gecko away from the vertical surface, while hindlegs produced forces that pulled the gecko toward the vertical surface. (E) Climbing geckos produced positive fore–aft forces that propelled the gecko upwards. (F) During climbing, all four legs pulled laterally towards the midline of the body such that the left legs produced forces that pulled the gecko to the left, while the right legs produced forces that pulled the gecko to the right. The directions of lateral GRFs during climbing were opposite to those produced during level running.





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