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Fig. 7. Schematic of proposed cross-linking in Nereis jaws. Zn-mediated
cross-linking of histidine-rich proteins is proposed to be responsible for
enhanced hardness and modulus in Nereis jaws. Cross-linking may be
intramolecular (black complexes) or intermolecular (grey complexes).
Additional cross-linking via di-and/or tri-tyrosine formation is not
expected to contribute significantly to mechanical properties of the bulk
jaw.