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Fig. 4. Schematic diagram illustrating two possible strategies of compensating
small-scale surface roughness (roughness smaller than the dimensions of the
pad). Good adhesive contact can either be achieved `dry' with very fine seta
tips or `wet' by secretion of a fluid that can fill out substrate cavities. A
fluid may not only be necessary in smooth adhesive pads (e.g. many insects and
tree frogs) as shown here but also in hairy pads, when setae are relatively
large and blunt as in many insects.