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Fig. 2. Recordings of CO2 emission from the anterior mesothoracic
spiracles and the posterior elytral case in four species of dung beetles. (A)
Pachylomerus femoralis (5.53 g), (B) Scarabaeus gariepinus
(1.475 g), (C) S. striatum (0.487 g) and (D) Circellium
bacchus (4.859 g). Note the different scales on the axes. C, closed
period; F, flutter period; B, burst period. These are explained further in the
text.