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Fig. 6. The `burst-and-glide' parameters R or Rp (A) and {alpha} (B) in relation to the average swimming speed of the great cormorant in each paddling cycle. Open symbols denote calculated values when the recovery is treated as part of the burst stage and full symbols are calculated values when the recovery is treated as part of the glide stage (see text). Solid horizontal lines mark the proportion of 1, where the energetic cost of burst-and-glide is exactly the same as for swimming at constant speed. Broken vertical lines in A mark the threshold average speed where the least square regression of the points (broken line for Rp, solid line for R) intersects with the value of 1. At this swimming speed the energetic benefit from burst-and-glide changes from gain (R<1) into loss (R>1). In B, values ≤1 imply that the drag coefficient for passive drag is as high as or higher than the coefficient for active swimming. Note that the open symbols represent a more conservative approach for estimating the energetic advantage from burst-and-glide.





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