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Fig. 5. Lateral projection of the geometric center of area of a Clione antarctica (individual no. 1) in vertical ascent during two complete wingbeat cycles. Arrows denote: a, the beginning of the power phase of the upstroke; b, recovery phase of the upstroke; c, power phase of downstroke; d, recovery phase of the downstroke; and e, the end of the stroke cycle.





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