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Fig. 6. Geometry of test space and outward journeys in experiment II. The nest box is located under the arena floor, the nest exit (diameter, 9 cm) being located at a distance of 50 cm from the arena wall. The hamster can easily climb from the underground nest into the arena. Before each test trial, the arena is rotated by 135°. The outward journey (continuous line with heavy arrows) leads from the (rotated) nest exit (N135°) over 60 cm to point P1, and, with the exception of fix-without-translation trials, over a further translation of 110 cm to point P2. In baseline trials, the animal proceeds to P1, where it takes up some food, and then to P2, where it fills its cheek pouches completely. Baseline-with-circling trials follow the same pattern except that the subject walks along two full circles near P1 (circle passing through P1) instead of taking up some food. In fix-without-translation trials, the animal walks to P1, follows two full circles in the dark, and then a further circle in the opposite direction under light. Fix-with-translation trials include an additional translation from point P1 to point P2 after the fix. During the fix at P1, the nest direction, as established by visual cues on the one hand and by PI on the other hand, diverge by ~67°. A full resetting would lead to the standard nest location N0°, and a partial (direction only) resetting to the virtual nest location NDR. The theoretical homing directions based on a full or a partial resetting differ by 72° at P1 and by 125° at P2.





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