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Fig. 3. Alterations in total lift and lift-to-drag ratio of the model fore- and hindwing in response to changes in kinematic phase shift between both wings. (A) Negative phase-shift values indicate that the forewing leads wing motion (red) throughout the entire stroke cycle whereas positive phase-shift values indicate that the hindwing leads wing motion (blue). Kinematic pattern of both wings is the same (cf. Fig. 4C). (B-D) Lift averaged over an entire stroke cycle (closed red circles) and lift-to-drag ratio (closed blue circles) are shown for the forewing flapping on top of the hindwing (B), the hindwing (C) and the combined performance of both fore- and hindwing (D) during various kinematic phase relationships. Performances of single wings flapping without wake-wing interaction are shown as solid lines in the respective color. Data are presented for the robotic wings vertically separated by 1.3 (closed circles) and 5.0 (open circles) mean forewing chord lengths, measured as the distance between the rotational axis of the two wings (cf. Fig. 1C). For an explanation of L, see text.





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