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Fig. 9. Modeled changes in mechanical work stroke–1 against drag, buoyancy, inertia (surge acceleration) and all three combined during ascent and horizontal swimming at the bottom (109 m), based on dive parameters for a Brünnich's guillemot (BRGU 13; Figs 3, 6 and 7; Table 1) and drag of a frozen Brünnich's guillemot (Fig. 1A). The depth of neutral buoyancy was estimated as 71 m (Fig. 3A). Values at the bottom were based on an estimated mean speed of 1.76 m s–1.





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