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Fig. 3. Free-flight flow visualization of the wake of the dragonfly Sympetrum sanguineum in counterstroking flight with the smoke-plane close to the right (far) wingtip. (A–J) Consecutive images from a 250 Hz high-speed video recording. In contrast to the centreline flow shown in Fig. 2, here the wingtip vortices are clear (purple arrows), and form wake elements (green arrows) that persist for several frames. However even here at the wingtips, where the wake structure is at its most coherent, the wake elements lose their identity after five frames (A–E, 1/50th of a second). The difference between the apparent structure of the wake elements between the tip region and the centreline region suggests that wake elements have a complex structure, consistent with the lack of any defined starting vortex.





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