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Fig. 1. Morphological phylogeny of the Family Scombridae showing its two
subfamilies, Gasterochismatinae and Scombrinae, the latter composed of four
Tribes. The bonitos (Tribe Sardini) and tunas (Thunnini) are sister groups.
Letter abbreviations for the five tuna genera: Al, Allothunnus;
A, Auxis; E, Euthynnus; K, Katsuwonus; T,
Thunnus. Modified from Collette et al.
(2001); Graham and Dickson
(2000).