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Fig. 4. The position of the digitized points representing the body of the great cormorant, relative to the mean swimming direction. (A) The center of mass of the bird demonstrated no significant deviations from the swimming direction in the Z direction (right/left). (B) Deviations in the Y direction (up/down) of three points on the body. Positive values are upward movements, and negative values are downward movements. The points are at anterior (base of neck), center (center of mass) and posterior (base of tail) locations on the body unit (points 2, 8 and 6 in Fig. 1C). All points descended during the stroke phase and ascended during the glide and recovery phases. The base of the neck (most anterior point) follows the pattern of the center of mass, indicating that the entire body is descending and ascending. The larger amplitude of the base of the tail can be attributed to the tilt of the body (rotation). The deviations are relative to the mean direction of the birds, calculated as the straight line connecting the position of point 3 in Fig. 1C (for the XY plane) or point 11 in Fig. 1B (for the XZ plane) at the beginning and end of the paddling cycle. The red broken arrows mark these directions. Each data point is the mean ± S.E.M. of all birds (N=9). The X axis is the same as described in Fig. 2.





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