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Fig. 1. Sketches of the model wings, the flapping motion and the reference frames. FW and HW denote fore- and hindwings, respectively. OXYZ is an inertial frame, with the X and Y axes in the horizontal plane; oxyz is another inertial frame, with the x and y axes in the stroke plane; o'x'y'z' is a frame fixed on the wing, with the x' axis along the wing chord and y' axis along the wing span. ß, stroke plane angle; {varphi}, positional angle; {alpha}, angle of attack; R, wing length.





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