Fig. 4. Endoscopic images illustrating a protrusion (pr) of the palatal organ during feeding on slurry off a sand substrate (500 frames s^-1; duration of sequence 64 ms; frames 1, 5, 8, 9, 10 and 12 shown). The anterior of the fish is to the right. Ceratobranchials I and II (cb I; cb II) are visible. (A) A protrusion of tissue from the palatal organ is projecting towards cb II. (B) The protrusion has come into contact with cb II. (C) A sand grain has entered the field of view on the left side of the protrusion during a flow reversal. (D) The sand grain passes lateral to the protrusion. (E) The protrusion begins to move dorsally and lifts from cb II while the water is still moving anteriorly during the flow reversal. (F) The palatal organ is returning to its original shape. Movie available online (movie: fig4.mov).

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