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Fig. 5. Summary of corpus and isthmus motions. For simulation, muscle motions were
assumed to be piecewise linear as diagrammed here (i.e. the extent of
contraction varies linearly with time, except at discrete breakpoints where
its rate of change with time abruptly changes.) The corpus contracts from 0 to
133 ms and relaxes from 133 to 150 ms. C indicates the fully contracted
position and R the fully relaxed position. The entire corpus contracts and
relaxes as a unit. The anterior isthmus contracts from 83 to 150 ms and
relaxes from 150 to 167 ms; the middle of the isthmus (the most posterior
region included in the simulation) contracts from 150 to 183 ms and relaxes
from 183 to 200 ms. Between the anterior and middle isthmus the times at which
contraction starts, relaxation starts, and relaxation ends vary linearly.
These times are based on the video sequence from which
Fig. 3 was abstracted, but the
precise times of pharyngeal motions are quite variable. Important features
consistently seen are the delay of isthmus motions with respect to the corpus,
and the progression of motion from the anterior to the middle isthmus.