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Fig. 1. (A) Schematic diagram of a hovering Drosophila showing the
orientation of the x,y,z coordinate system. (B) Schematic diagram of
the flapping Drosophila wing. The position of the wing is shown at
three different times during the flapping cycle. The coordinate system
(x',y',z') is fixed to the wing, and the wing
rotates about the z' axis throughout the cycle. R,
wing length; 
, wingbeat amplitude.
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