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Fig. 2. Example time series of forces applied to seaweeds in the field. (A) Bending moments (i.e. forces applied with a given lever arm) imposed on the base of the stipe of an upright understory kelp Pterygophora californica exposed to unbroken waves. Note the relatively regular character of the recording. (B) Tensile force in a stipe of Nereocystis luetkeana exposed to unbroken waves. The slightly more complicated trajectory of force (which varies around that induced by the plant's buoyancy alone) probably arises from the seaweed's interaction with the water's surface as it is swept back and forth by drag. (C) A shorter time series of force applied to an emergent intertidal alga, Pelvetia compressa. The far more complicated fluid motions associated with wave impingement and breaking produce a rapidly evolving force record, particularly at the instant of wave arrival. Note that the magnitude of forces imposed on this small surfzone plant (frond length 15 cm) rival those acting on the far larger (7 m length) subtidal bull kelp (compare B and C). This is a direct consequence of the greater severity of flow in intertidal regions.





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