First published online November 1, 2006
Journal of Experimental Biology 209, 4566-4573 (2006)
Published by The Company of Biologists 2006
doi: 10.1242/jeb.02536
Seasonal adjustment of energy budget in a large wild mammal, the Przewalski horse (Equus ferus przewalskii) II. Energy expenditure
Walter Arnold*,
Thomas Ruf and
Regina Kuntz
Research Institute of Wildlife Ecology, University of Veterinary
Medicine, Vienna, Savoyenstraße 1, 1160 Vienna, Austria

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Fig. 1. Seasonal changes in physiological parameters and behaviour (daily means;
measures of variation reflect differences between individuals). Data are
double plotted to ease visualization of the annual pattern. Labels on the
x-axis indicate the first month of a two-month interval. Only
complete data sets, i.e. those with information about all parameters are
plotted. (A) Daily means of heart rate (means ± s.e.m.). (B) Daily
means of locomotor activity (means ± s.e.m.). (C) Top: daily means and
range of subcutaneous temperature; bottom: daily means of air temperature.
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Fig. 2. Daily mean heart rate of three pregnant mares from 1 March 2003 to 31
August 2003. Different symbols represent different individuals, arrows
represent the time of parturitions.
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Fig. 3. Daily mean heart rate during locomotor activity (simultaneously measured
locomotor activity >10%; open circles, means + s.e.m.) and at rest
(locomotor activity 10%; closed circles, means - s.e.m.).
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Fig. 4. Upper graph: daily mean heart rate (closed circles) during or close to the
day of measurement of heat increment of feeding. Lower graph: estimate of the
heat increment of feeding (open circles) and its seasonal change calculated
from data presented in Kuntz et al. (Kuntz
et al., 2006 ). Lines represent periodic fits to the data, vertical
lines represent the acrophases.
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© The Company of Biologists Ltd 2006