First published online November 17, 2005
Journal of Experimental Biology 208, 4419-4426 (2005)
Published by The Company of Biologists 2005
doi: 10.1242/jeb.01909
An experimental study of social attraction and spacing between the sexes in sheep
Pablo Michelena1,*,
Karine Henric1,
Jean-Marc Angibault2,
Jacques Gautrais1,
Paul Lapeyronie3,
Richard H. Porter4,
Jean-Louis Deneubourg5 and
Richard Bon1
1 Centre de Recherches sur la Cognition Animale, Université Paul
Sabatier, F-31062, France
2 Laboratoire de Comportement et d'Ecologie de la Faune Sauvage, INRA, 31326
Castanet-Tolosan cedex, France
3 UMR Elevage des Ruminants en Région Chaude, Agro-M-INRA-CIRAD,
34060 Montpellier Cedex 1, France
4 Laboratoire de Comportement, INRA-CNRS, 37380 Nouzilly, France
5 Service d'Ecologie Sociale, Université Libre de Bruxelles,
Bruxelles

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Fig. 1. Experimental design used to measure the level of attraction exerted by one
or two stimulus conspecifics. R, releasing cage; S, stimulus cage. During the
30 min experiments, snapshots were taken at 1 min intervals using a digital
camera anchored on the tower.
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Fig. 2. Distribution of the observed (bars) and expected (horizontal line)
probability of presence in 2 m rings of test males and females as a function
of the distance to the cages. The observed probability is the proportion of
the locations/2 m ring surface. The expected probability is calculated under
the hypothesis of random distribution of sheep. Gray bars represent the
distributions relative to the same-sex (B) and opposite-sex stimulus (C) and
white bars to the empty cage. When confronted with two empty cages (A) or two
same-sex stimuli (D) the grey and white bars refer, respectively, to the right
and left cages. With two opposite-sex stimuli (E), grey bars represent the
distribution relative to the stimulus of same sex as the test sheep, and white
bars to the opposite sex.
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© The Company of Biologists Ltd 2005