First published online July 6, 2005
Journal of Experimental Biology 208, 2625-2631 (2005)
Published by The Company of Biologists 2005
doi: 10.1242/jeb.01609
Running performance has a structural basis
Peter G. Weyand1,2,* and
J. Adam Davis1
1 Locomotion Laboratory, Kinesiology Department, MS545, Rice
University, 6100 Main Street, Houston, TX 77005, USA
2 Center for Human Performance, Texas Medical Center, Houston, TX 77030,
USA

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Fig. 1. Mass-specific stance-averaged support forces vs running speed. (A)
stance-averaged force vs running speed in a representative subject;
(B) mean stance-averaged support forces for athletic male (M; N=9)
and female (F; N=9) subjects (M: y=1.27 + 0.105x;
F: y=1.24 + 0.106x). Vertical hashes across the horizontal
lines appearing in the upper portion represent the average race velocities of
elite male and female specialists at each event distance. y-intercept
values of Fg typically do not equal 1.0 because they are
extrapolations beyond the range of speeds at which running is possible.
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Fig. 2. Body sizes of elite specialists vs event distance (means ±
S.E.M.). (A) Body mass; (B) height.
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Fig. 3. Body mass index (BMI) and ground support forces vs event distance.
(A) Body mass index; (B) ground support forces determined from the average
racing speeds of event specialists; (C) the ratio of BMI to mass-specific
ground support force of specialists vs event distance (mean BMI
Fg1=9.92 kg m2; broken
horizontal line).
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Fig. 4. The product of ground support force and height-squared in relation to body
mass (mass=0.006+0.101x, R2=0.97).
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© The Company of Biologists Ltd 2005