First published online December 22, 2003
Journal of Experimental Biology 207, 449-460 (2004)
Published by The Company of Biologists 2004
doi: 10.1242/jeb.00739
Unsteady forces and flows in low Reynolds number hovering flight: two-dimensional computations vs robotic wing experiments
Z. Jane Wang1,*,
James M. Birch2 and
Michael H. Dickinson3
1 Theoretical and Applied Mechanics, Cornell University, Ithaca, NY 14853,
USA
2 Integrative Biology, University of California, Berkeley, CA 94720,
USA
3 Bioengineering, California Institute of Technology, Pasadena, CA 91125,
USA

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Fig. 6. Vorticity plot in the case of A0/c=4.8,
=0. Ten frames are shown in the fourth stroke. Red, counterclockwise
rotating vortices; blue, clockwise rotating vortices. The wing is in black.
(A,C) Computed vorticity; (B,D) digital particle image vorticity data in a 2D
slice at 0.65R. See Materials and methods for details. Each pair in
A,B and C,D corresponds to the same time during a stroke. The time sequence is
indicated by the numbers on each plate. The color scale for vorticity of
computation and experiments do not correspond to the exact same contour
values, so the figure should be viewed as a qualitative comparison.
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Fig. 1. Quasi-steady lift CL (circles) and drag
CD (crosses) coefficients measured from computation
compared to the empirical formulae described by Equations
14,15
(solid and broken lines, respectively).
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Fig. 3. Computational and experimental lift and drag coefficients during
symmetrical rotation ( =0; A0/c=2.8).
Details as in Fig. 2.
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Fig. 7. Force comparison between experiment, computation and quasi-steady
predictions for (A,D) advanced ( = /4), (B,E) symmetric ( =0) and
(C,F) delayed ( = /4) rotations. (AC) Force traces of
four strokes starting from rest. Red, experimental force; blue, computational
force; green, quasi-steady estimates using Equations
14,
15,
16,
17. Forces are normalized as in
Figs 2,
3,
4. (DF) Difference trace
(red) between the experimental forces and quasi-steady forces. Linear
acceleration du(t)/dt (blue), and an estimate of
rotational force u(t) (t) (green) are shown
in arbitrary scale since we are only interested in their basic time course.
Broken vertical lines mark the wing reversal during t=[1,2]. Features
associated with rotation (r) and unsteady circulation due to wing acceleration
(u), are labelled accordingly.
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© The Company of Biologists Ltd 2004