First published online April 8, 2004
Journal of Experimental Biology 207, 1601-1606 (2004)
Published by The Company of Biologists 2004
doi: 10.1242/jeb.00908
Paradox lost: answers and questions about walking on water
Mark W. Denny
Hopkins Marine Station, Stanford University, Pacific Grove, CA
93950-3094, USA

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Fig. 1. A water strider standing motionless on the water's surface. Note the
dimples where the feet contact the water.
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Fig. 2. Surface tension (which acts parallel to the airwater interface)
pulls upward on the leg of a water strider.
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Fig. 3. The speed of a pure capillary wave decreases with an increase in
wavelength, while the speed of a pure gravity wave increases with an increase
in wavelength. For real waves, the result is a minimum wave speed.
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Fig. 4. As a water strider sweeps its middle legs backward, momentum is produced in
the water associated with hemispherical vortices.
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© The Company of Biologists Ltd 2004