Fig. 4. The evolution of locomotor biomechanics in relation to foraging mode in lizards. (A) Ancestral character reconstruction of phase shift values (from B) are mapped onto the lizard phylogeny from Townsend et al. (Townsend et al., 2004) in A. Branch shading indicates running (black), walking (white), and the grey branches leading to the base of the Gekkota and the Scincidae are equivocal for running or walking. (B) Raw phase shift data indicating running (closed symbols, phase shift 45) and walking (open symbols, phase shift 135) mechanics. (C) Reconstructed patterns of foraging mode evolution from Fig. 1 (black branches are SW lineages, white branches are WF lineages, hatched branch is unknown). Note that walking mechanics evolved each time WF evolved. Asterisks indicate species taken from the literature.