Fig. 13. Summary of the evolution of Tadros' morphology, mechanics and behavior. Plus and minus signs represent a statistically significant (P<0.05) correlation or regression. Blue arrows indicate relations established in biomechanical analysis (Fig. 9). Red arrow indicates relation detected only in competition trials (Fig. 8). Green arrows indicate relations by formulaic definition (Eqn 1, Eqn 3). Broken lines with double-headed arrows indicate correlations among kinematic phenotypes during competition (Table 4). Solid black lines with single-headed arrows show conceptual path of the phenotypic system through the genotypic manipulations that produce novel offspring from the adult population.