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Fig. 2. Recordings of CO2 emission from the anterior mesothoracic spiracles and the posterior elytral case in four species of dung beetles. (A) Pachylomerus femoralis (5.53 g), (B) Scarabaeus gariepinus (1.475 g), (C) S. striatum (0.487 g) and (D) Circellium bacchus (4.859 g). Note the different scales on the axes. C, closed period; F, flutter period; B, burst period. These are explained further in the text.