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Fig. 5. Comparison of landmark fidelity, i.e. the tendency for bees to stay faithful to the landmark during search, with odometry (black line) and without odometry (grey line). Bees with odometry remained faithful on all four {cup}-turns, whereas bees without odometry progressively gave up searching at the landmark, in all conditions. (A) Landmark in place at the training site, (B) landmark shifted to unit 14, and (C) landmark shifted to unit 4. Numbers given indicate the number of bees considered when calculating the observed frequency.