Fig. 2. Results of Experiment 1. (A) Bees tested with no landmark conducted a broad search. (B) With the landmark at the training location, bees searched very accurately. (C) When the landmark was shifted to unit 14, bees searched either at the landmark or the training location. (D) Bees searched at the landmark when it was shifted to unit 4. Black bars, first -turns; grey bars, second -turns; coloured lines, search distributions; inverted triangles, training location; diamonds, landmark position; N=flight number. Note different y-axis scales in A,C and B,D.