Fig. 1. Cylindrical flight chamber (height and diameter, 2.4 m). The circular floor is filmed from above using a video camera (VC). Bees entered the tent through the entry tube (ET), attached to a white painted side wall (SW). Bees flew to the 1.9 m-distant food hole (FH), which gave access to a feeder (not shown). Discs on the floor provided visual feedback. The inset shows a plan view of the set-up with the locations of the landmarks used in the experiments: (1) paper square (side length, 25 cm); (2, 3) cylinder (diameter and height, 25 cm) located 0.5 m to the side of the food hole; (4) same cylinder, located 1 m to the right of the center.