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Fig. 4. (A) Test of E. lucunter with two spines. Off-angle light (gray arrows) is absorbed and reflected by the spines. Light within the acceptance angle, {theta} (white arrow), reaches the body wall between the spines and is detected. (B) E. lucunter in a typical shelter. (C) View of B at a spatial resolution of 33°. Scale bars are 20°, which implies that the center of the viewing echinoid is approximately 6 cm from the center of the pictured echinoid.