Fig. 3. Tilting angles of the body, tail and neck (_B, _T and _N, respectively) during the paddling cycle of a swimming great cormorant. Angles are relative to the mean swimming direction (see text). The angles are 0 throughout the paddling cycle. _B and _T are in anti-phase during the stroke phase, and _T remains at a steeper angle than _B during the glide and recovery phases. Each data point on the curve is the mean ± S.E.M. of all birds (N=9). The X axis is the same as described in Fig. 2 and in the text.