Fig. 1. Tongue aiming ability was quantified by having individuals of Phrynomantis bifasciatus aim into five quadrants: (A) left –46° to –105°, (B) left –6° to –45°, (C) 0° to 5° to either side, (D) right 6° to 45°, (E) right 46° to 105°. The quadrant is essentially a bib, with the midline of the head designating 0°. As the head of the animal turns, the quadrant follows this movement so that a line drawn down the midline of the head would always be located at 0°.