Fig. 1. (A) Water loss rate (mg h^-1) adjusted for body mass, and (B) proportional contribution of spiracular to total water loss (%), adjusted for body mass, at 20°C in the five Scarabaeus species investigated in this study. Variation in rate of water loss is in the direction expected from the mean annual rainfall of the area from which the species was collected. Values are means ± S.E.M. Gari, S. gariepinus; Stri, S. striatus; Gale, S. galenus; Rust, S. rusticus; West, S. westwoodi.