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Fig. 1. Schematic of the force balance for the drag measurements. The crab was supported at a distance l1 below the pivot point. A weight (W) on the horizontal lever arm, at a distance l2 from the pivot point, provided a moment in the opposite sense to that of the drag force (FD). Fshaft represents the drag force on the submerged section of the support shaft, which is at a distance lshaft below the pivot point.