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Fig. 1. (A) Diagram of the fly and host defining the variables used for the analysis. The angle of the fly’s longitudinal axis ({Phi}) is measured relative to externally based coordinates. The angular position of the host head ({theta}H) and tail ({theta}T) are measured relative to {Phi}. (B) Diagram of the fly in two frames to illustrate the fly’s movement, consisting of rotation ({Delta}{Phi}) and translation. Translation is resolved into two components, forward (F) and sideways (S).