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Fig. 4. The relationship between anterior and posterior CO2 emission rates during the discontinuous gas-exchange cycle (DGC) in Circellium bacchus (N=53 DGCs in six beetles, mean mass 7.427 g). The regression equations are: a=-0.022+0.891t, r2=0.98, P=0.001; p=0.022+0.109t, r2=0.4, P=0.001, where a is anterior spiracular CO2, p is posterior spiracular CO2 and t is total CO2. When the mean values for each beetle (N=6) are used, the regression equations are: a=-0.021+0.866t, r2=0.98, P=0.002; p=0.021+0.134t, r2=0.5, P=0.11.