Fig. 5. Examples of searching trajectories for two ants recorded after releases from four different compass directions after a short training period (see text). Open black circles indicate the positions of the landmarks; stars represent the sites of release. The green lines indicate the search trajectories of ants released from southeast, i.e. from the direction from which they had approached the nest during training. According to the criteria used to define the number of search positions, ants 235 and 311 both exhibited four search positions.